Redescription of Ellimma branneri and Diplomystus shengliensis, and relationships of some basal clupeomorphs

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	Redescription of Ellimma branneri and Diplomystus shengliensis, and relationships of some basal clupeomorphs
	Chang, MM ; Maisey, JG ; Chang, MM (reprint author), Amer Museum Nat Hist, Div Paleontol, New York, NY 10024 USA.
	2003-05-22
发表期刊	AMERICAN MUSEUM NOVITATES
ISSN	0003-0082
期号	3404 页码:1-35
文章类型	Article
摘要	Two extinct clupeomorphs, daggerEllimma branneri from the Cretaceous of Brazil and daggerDiplomystus shengliensis from the Eocene of China, are redescribed. daggerEllimma branneri was formerly classified within the Clupeiformes, but it lacks derived characters of clupeiforms and clupeoids. Dorsal scute "wings" are expanded and subrectangular in daggerEllimma and other members of the family daggerParaclupeidae Chang and Chou (1977), approximately equal to daggerEllimmichthyidae Grande (1982a). Consequently, daggerEllimma branneri is classified here within the family daggerParaclupeidae. daggerParaclupeidae are known from the Lower Cretaceous to the middle Eocene. In the present work, two monophyletic groups are identified within the daggerParaclupeidae. One group (subfamily daggerParaclupeinae of Chang and Grande, 1997), known only from the Lower Cretaceous (Hauterivian-Albian), includes daggerParaclupea, daggerEllimmichthys, and daggerEllimma. These taxa are united by strongly sculptured, skull-roofing bones with ridges radiating from the growth center, and a dorsal scute ornament of prominent ridges. daggerScutatuspinosus may also belong in this group. The other group includes daggerDiplomystus (Upper Cretaceous-Eocene) and daggerArmigatus (Upper Cretaceous), which are united by a single homoplaseous character (presence of a posteriorly expanded third hypural, leaving no gap between hypurals 2 and 4): this character also occurs in pristigasteroids, daggerErichalcis, osteoglossids, some elopomorphs (daggerLe-banichthys lewisi, and most Albula spp.), and a number of ostariophysans not included in our analysis. daggerParaclupeines are customarily regarded as being more closely related to the Clupeiformes than to other teleosts (i.e., as clupeomorphs), although no derived characters are uniquely shared by daggerEllimma branneri and modern Clupeiformes. The relationships of daggerEllimma and certain other extinct herring-like teleosts (including other daggerparaclupeines) with the Clupeiformes are unclear, and they may collectively form a paraphyletic assemblage. No biogeographical hypothesis satisfactorily explains the known distribution of nonmarine daggerparaclupeine fishes in the Cretaceous. A substantial portion of their nonmarine fossil record is missing (as evidenced by the recent discovery of a possible daggerparaclupeine, daggerEzkutuberezi carmeni Poyato-Ariza et al., 2000, in Spain), and some aspects of their early distribution pattern may have involved marine dispersal. Eocene daggerDiplomystus occurs on both sides of the Pacific Ocean, but the "Pacifica" hypothesis (which lacks empirical support) is abandoned as an explanation for such Eocene (and younger) trans-Pacific distribution patterns of nonmarine fishes. Instead, a "freshwater Arctic Ocean" hypothesis is favored. According to this hypothesis (for which there are several independent lines of geological evidence), temporary desalination of the Arctic Ocean occurred during the Paleocene and early Eocene, which may have permitted freshwater fishes to move unimpeded by salt-water barriers between Asia and North America; this temporary desalination event may eventually become recognized as a significant factor in the holarctic distribution patterns of various Tertiary-Recent freshwater fishes.; Two extinct clupeomorphs, daggerEllimma branneri from the Cretaceous of Brazil and daggerDiplomystus shengliensis from the Eocene of China, are redescribed. daggerEllimma branneri was formerly classified within the Clupeiformes, but it lacks derived characters of clupeiforms and clupeoids. Dorsal scute "wings" are expanded and subrectangular in daggerEllimma and other members of the family daggerParaclupeidae Chang and Chou (1977), approximately equal to daggerEllimmichthyidae Grande (1982a). Consequently, daggerEllimma branneri is classified here within the family daggerParaclupeidae. daggerParaclupeidae are known from the Lower Cretaceous to the middle Eocene.
WOS标题词	Science & Technology ; Life Sciences & Biomedicine
URL	查看原文
关键词[WOS]	ELLESMERE ISLAND ; EOCENE ; FISH ; MAMMALS ; CHINA
收录类别	SCI
语种	英语
WOS研究方向	Biodiversity & Conservation ; Zoology
WOS类目	Biodiversity Conservation ; Zoology
WOS记录号	WOS:000183145700001
引用统计
文献类型	期刊论文
条目标识符	http://119.78.100.205/handle/311034/3727
专题	中科院古脊椎所（2000年以后）
通讯作者	Chang, MM (reprint author), Amer Museum Nat Hist, Div Paleontol, New York, NY 10024 USA.
作者单位	1.Amer Museum Nat Hist, Div Paleontol, New York, NY 10024 USA 2.Chinese Acad Sci, Inst Vertebrate Paleontol & Paleoanthropol, Beijing 100044, Peoples R China
推荐引用方式 GB/T 7714	Chang, MM,Maisey, JG,Chang, MM . Redescription of Ellimma branneri and Diplomystus shengliensis, and relationships of some basal clupeomorphs[J]. AMERICAN MUSEUM NOVITATES,2003(3404):1-35.
APA	Chang, MM,Maisey, JG,&Chang, MM .(2003).Redescription of Ellimma branneri and Diplomystus shengliensis, and relationships of some basal clupeomorphs.AMERICAN MUSEUM NOVITATES(3404),1-35.
MLA	Chang, MM,et al."Redescription of Ellimma branneri and Diplomystus shengliensis, and relationships of some basal clupeomorphs".AMERICAN MUSEUM NOVITATES .3404(2003):1-35.

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